The marginal wetlands that remain continue to be threatened by reclamation for farms and homes.

怎么樣理解此處的remain continue？難道remain后面可以接動詞嗎？that remain不是做The marginal wetlands謂語嗎？continue這里又是一個謂語？雙謂語嗎？

The marginal wetlands that remain continue to be threatened by reclamation for farms and homes.that remain是定語從句，謂語是continue。