An international team of scientists has successfully decoded ancient DNA from animal skin on which some of the Dead Sea Scrolls were written. 在這個句子on which some of the Dead Sea Scrolls中on which, some of the Dead Sea Scrolls是什么關系？在這個句子on which some of the Dead Sea Scrolls were written.中哪個是主語，謂語，定語？

on which some of the Dead Sea Scrolls were written為定語從句，修飾ancient DNA from animal skin。

定語從句的主語為some of the Dead Sea Scrolls 謂語動詞為were written, 狀語為on which. on=about. which指代ancient DNA from animal skin.